About the Authors

Mohd Zamri Bin Haji Ismail

Contributed equally to this work with: Mohd Zamri Bin Haji Ismail, Matt D. Hodges

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Matt D. Hodges

Contributed equally to this work with: Mohd Zamri Bin Haji Ismail, Matt D. Hodges

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Michael Boylan

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Rajesh Achall

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Alan Shirras

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Susan J. Broughton

s.j.broughton@lancaster.ac.uk

Affiliation Division of Biomedical and Life Sciences, Faculty of Health and Medicine, Lancaster University, Lancaster, LA1 4YQ, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJB. Performed the experiments: SJB MZBHI MDH MB RA. Analyzed the data: SJB MZBHI MDH AS. Contributed reagents/materials/analysis tools: AS. Wrote the paper: SJB.