About the Authors

Kazumichi Nagasawa

Affiliation Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Mizue Meguro

Affiliation Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Kei Sato

Affiliation Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Yuta Tanizaki

Affiliations Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan, Department of Biology, School of Education, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Nami Nogawa-Kosaka

Affiliation Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Takashi Kato

tkato@waseda.jp

Affiliations Integrative Bioscience and Biomedical Engineering, Graduate School of Advanced Science and Engineering, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan, Department of Biology, School of Education, Waseda University, Center for Advanced Biomedical Science, TWIns building, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KN NNK TK. Performed the experiments: KN. Analyzed the data: KN MM KS YT TK. Wrote the paper: KN TK.