About the Authors

Heather R. Conti

Affiliation University of Pittsburgh, Department of Medicine, Division of Rheumatology & Clinical Immunology, Pittsburgh, PA, United States of America

Natasha Whibley

Affiliation University of Pittsburgh, Department of Medicine, Division of Rheumatology & Clinical Immunology, Pittsburgh, PA, United States of America

Bianca M. Coleman

Affiliation University of Pittsburgh, Department of Medicine, Division of Rheumatology & Clinical Immunology, Pittsburgh, PA, United States of America

Abhishek V. Garg

Affiliation University of Pittsburgh, Department of Medicine, Division of Rheumatology & Clinical Immunology, Pittsburgh, PA, United States of America

Jillian R. Jaycox

Affiliation Carnegie Mellon University, Dept. of Biological Sciences, Pittsburgh, PA, United States of America

Sarah L. Gaffen

sarah.gaffen@pitt.edu

Affiliation University of Pittsburgh, Department of Medicine, Division of Rheumatology & Clinical Immunology, Pittsburgh, PA, United States of America

Competing Interests

SLG has received a research grant, travel reimbursements and honoraria from Novartis. SLG has also consulted for, received travel reimbursements and honoraria from Janssen, Eli Lilly, Amgen and Pfizer. There are no other conflicts of interest. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SLG HRC NW. Performed the experiments: HRC NW BMC AVG JRJ. Contributed reagents/materials/analysis tools: SLG HRC. Wrote the paper: HRC SLG.