About the Authors

Benjamin J. van Soldt

Affiliation Institute of Biology, University of Leiden, Leiden, the Netherlands

Brian D. Metscher

Affiliation Department of Theoretical Biology, University of Vienna, Vienna, Austria

Robert E. Poelmann

Affiliation Department of Anatomy and Embryology, Leiden University Medical Center, Leiden, the Netherlands

Bart Vervust

Affiliation Department of Biology, University of Antwerp, Antwerp, Belgium

Freek J. Vonk

Affiliations Institute of Biology, University of Leiden, Leiden, the Netherlands, NCB Naturalis, Leiden, the Netherlands

Gerd B. Müller

Affiliation Department of Theoretical Biology, University of Vienna, Vienna, Austria

Michael K. Richardson

m.k.richardson@biology.leidenuniv.nl

Affiliation Institute of Biology, University of Leiden, Leiden, the Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BJVS REP MKR. Performed the experiments: BJVS BDM. Analyzed the data: BJVS REP MKR. Contributed reagents/materials/analysis tools: BDM GM BV FJV MKR. Wrote the paper: BJVS MKR.