About the Authors

Gang Liu

‡ GL and GY contributed equally to this paper.

Affiliation Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China

Guan Yang

‡ GL and GY contributed equally to this paper.

Affiliation College of Animal science & Technology, Hunan Agricultural University, 410128, Changsha, Hunan, China

Guiping Guan

Affiliation College of Animal science & Technology, Hunan Agricultural University, 410128, Changsha, Hunan, China

Yuzhe Zhang

yinyulong@isa.ac.cn (YY); 18988669786@189.cn (YZ)

Affiliation Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China

Wenkai Ren

Affiliations Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China, Graduate University of the Chinese Academy of Sciences, Beijing, 100039, China

Jie Yin

Affiliations Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China, Graduate University of the Chinese Academy of Sciences, Beijing, 100039, China

Yordan Martínez Aguilar

Affiliation Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China

Wei Luo

Affiliation College of Veterinarian, Hunan Agricultural University, Changsha, Hunan, 410128, China

Jun Fang

Affiliation College of Animal science & Technology, Hunan Agricultural University, 410128, Changsha, Hunan, China

Xinglong Yu

Affiliation College of Veterinarian, Hunan Agricultural University, Changsha, Hunan, 410128, China

Tiejun Li

Affiliation Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China

Yulong Yin

yinyulong@isa.ac.cn (YY); 18988669786@189.cn (YZ)

Affiliation Scientific Observing and Experimental Station of Animal Nutrition and Feed Science in South-Central, Ministry of Agriculture, Hunan Provincial Engineering Research Center of Healthy Livestock and Poultry, Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, Hunan, 410125, China

Competing Interests

The authors have declared that no competing interests exist. The authors also declare the affiliations to this company, along with any other relevant declarations relating to employment, consultancy, patents, products in development or marketed products, etc. The authors also confirm that: This does not alter the authors’ adherence to PLOS ONE polices on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GL GY YY. Performed the experiments: GL GY GG JY WR YMA. Analyzed the data: WL JF XY TL. Contributed reagents/materials/analysis tools: YZ XY TL YY. Wrote the paper: GL GY WR.