About the Authors

Kazuhiro Chiku

Contributed equally to this work with: Kazuhiro Chiku, Takanori Nihira

Affiliation Faculty of Agriculture, Niigata University, Niigata, Japan

Takanori Nihira

Contributed equally to this work with: Kazuhiro Chiku, Takanori Nihira

Affiliation Faculty of Agriculture, Niigata University, Niigata, Japan

Erika Suzuki

Affiliation Faculty of Agriculture, Niigata University, Niigata, Japan

Mamoru Nishimoto

Affiliation National Food Research Institute, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Motomitsu Kitaoka

Affiliation National Food Research Institute, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Ken'ichi Ohtsubo

Affiliation Faculty of Agriculture, Niigata University, Niigata, Japan

Hiroyuki Nakai

nakai@agr.niigata-u.ac.jp

Affiliation Faculty of Agriculture, Niigata University, Niigata, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MK HN. Performed the experiments: KC TN ES. Analyzed the data: KC TN ES MK HN. Contributed reagents/materials/analysis tools: MN KO MK. Wrote the paper: KC MK HN.