About the Authors

Tim K. Davies

t.davies@mrag.co.uk

Affiliation MRAG Ltd, London, United Kingdom

Chris C. Mees

Affiliation MRAG Ltd, London, United Kingdom

E. J. Milner-Gulland

Affiliation Imperial College London, London, United Kingdom

Competing Interests

Two of the authors, Tim K. Davies and Chris C. Mees, are currently employed at a commercial company (MRAG Ltd). This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TKD EJMG CCM. Performed the experiments: TKD. Analyzed the data: TKD. Contributed reagents/materials/analysis tools: TKD. Wrote the paper: TKD.