About the Authors

Claire E. Stewart

Affiliation School of Biology, University of St Andrews, Fife, Scotland, United Kingdom

Richard E. Randall

Affiliation School of Biology, University of St Andrews, Fife, Scotland, United Kingdom

Catherine S. Adamson

csa21@st-andrews.ac.uk

Affiliation School of Biology, University of St Andrews, Fife, Scotland, United Kingdom

Competing Interests

UK Patent Application No. 1310432.8 in the name of University Court of the University of St Andrews. Patent Application No. PCTGB2014/051812, Title: Virus Production. This does not alter the authors' adherence to all PLOS ONE policies on data sharing or material availability.

Author Contributions

Conceived and designed the experiments: CSA RER CES. Performed the experiments: CSA CSE. Analyzed the data: CSA CES RER. Contributed reagents/materials/analysis tools: CSA RER. Wrote the paper: CSA RER.