About the Authors

Hayato Hirai

Affiliation Laboratory of Cytoskeletal Logistics, Department of Life Science and Medical Bioscience, Graduate School of Advanced Science and Technology, Waseda University, TWIns, Tokyo, Japan

Kunio Arai

Affiliations Laboratory of Cytoskeletal Logistics, Department of Life Science and Medical Bioscience, Graduate School of Advanced Science and Technology, Waseda University, TWIns, Tokyo, Japan, Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Tokyo, Japan

Ryo Kariyazono

Affiliation Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Tokyo, Japan

Masayuki Yamamoto

Affiliations Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Tokyo, Japan, Laboratory of Cell Responses, National Institute for Basic Biology, Aichi, Japan

Masamitsu Sato

masasato@waseda.jp

Affiliations Laboratory of Cytoskeletal Logistics, Department of Life Science and Medical Bioscience, Graduate School of Advanced Science and Technology, Waseda University, TWIns, Tokyo, Japan, Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Tokyo, Japan, PRESTO, Japan Science and Technology Agency, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KA MS. Performed the experiments: HH KA RK. Analyzed the data: HH KA RK. Contributed reagents/materials/analysis tools: HH KA RK MS. Wrote the paper: KA MS. Supervision: MY.