About the Authors

Raul J. Cano

rcano@calpoly.edu

Affiliation Center for Applications in Biotechnology, California Polytechnic State University, San Luis Obispo, California, United States of America

Jessica Rivera-Perez

Affiliation Department of Biology, University of Puerto Rico, San Juan, Puerto Rico

Gary A. Toranzos

Affiliation Department of Biology, University of Puerto Rico, San Juan, Puerto Rico

Tasha M. Santiago-Rodriguez

Affiliation Department of Pathology, University of California San Diego, San Diego, California, United States of America

Yvonne M. Narganes-Storde

Affiliation Center for Archaeological Research, University of Puerto Rico, Rio Piedras Campus, San Juan, Puerto Rico

Luis Chanlatte-Baik

Affiliation Center for Archaeological Research, University of Puerto Rico, Rio Piedras Campus, San Juan, Puerto Rico

Erileen García-Roldán

Affiliation Department of Biology, University of Puerto Rico, San Juan, Puerto Rico

Lucy Bunkley-Williams

Affiliation Department of Biology, University of Puerto Rico, Mayaguez Campus, San Juan, Puerto Rico

Steven E. Massey

Affiliation Department of Biology, University of Puerto Rico, San Juan, Puerto Rico

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJC SEM. Performed the experiments: RJC JRP EGR YNS LCB. Analyzed the data: RJC SEM GAT YNS TSR LBW. Contributed reagents/materials/analysis tools: RJC SEM GAT. Contributed to the writing of the manuscript: RJC SEM TSR JRP. Obtained permissions for collecting coprolites: YNS LCB.