About the Authors

Yusuke Tsujimura

Affiliation Laboratory of Immunoregulation and Vaccine Research, Tsukuba Primate Research Center, National Institute of Biomedical Innovation, Tsukuba, Ibaraki, Japan

Hiroyasu Inada

Affiliation Department of Pharmaceutical Sciences, Suzuka University of Medical Science, Suzuka, Mie, Japan

Misao Yoneda

Affiliation Department of Pathologic Oncology, Institute of Molecular and Experimental Medicine, Faculty of Medicine, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Tomoyuki Fujita

Affiliation Research Laboratories, Kyoto R&D Center, Maruho Co., Ltd, Chudoji, Shimogyo-ku, Kyoto, Japan

Kazuhiro Matsuo

Affiliation Research and Development Department, Japan BCG Laboratory, Kiyose, Tokyo, Japan

Yasuhiro Yasutomi

yasutomi@nibio.go.jp

Affiliations Laboratory of Immunoregulation and Vaccine Research, Tsukuba Primate Research Center, National Institute of Biomedical Innovation, Tsukuba, Ibaraki, Japan, Department of Immunoregulation, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Competing Interests

The authors have no commercial or financial conflict of interest. The authors also have no competing interest in Maruho Co., Ltd. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YY. Performed the experiments: YT HI MY. Analyzed the data: YY KM. Contributed reagents/materials/analysis tools: TF. Contributed to the writing of the manuscript: YT YY.