About the Authors

Lisa E. Wolber

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Claire J. Steves

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Pei-Chien Tsai

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Panos Deloukas

Affiliations William Harvey Research Institute, Barts and The London School of Medicine and Dentistry, Queen Mary University of London, London, United Kingdom, King Abdulaziz University, Jeddah, Saudi Arabia, Wellcome Trust Sanger Institute, Hinxton, Cambridge, United Kingdom

Tim D. Spector

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Jordana T. Bell

Contributed equally to this work with: Jordana T. Bell, Frances M. K. Williams

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Frances M. K. Williams

Contributed equally to this work with: Jordana T. Bell, Frances M. K. Williams

frances.williams@kcl.ac.uk

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LEW JTB FMKW PT CJS. Performed the experiments: LEW PT CJS. Analyzed the data: LEW. Contributed reagents/materials/analysis tools: PD TDS CJS. Wrote the paper: LEW.