About the Authors

John V. Roughan

j.v.roughan@ncl.ac.uk

Affiliation Comparative Biology Centre, The Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom

Claire A. Coulter

Affiliation Comparative Biology Centre, The Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom

Paul A. Flecknell

Affiliation Comparative Biology Centre, The Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom

Huw D. Thomas

Affiliation Northern Institute for Cancer Research, The Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom

Kenneth J. Sufka

Affiliation Research Institute of Pharmaceutical Sciences and Departments of Psychology and Pharmacology, Peabody Building, University of Mississippi, Oxford, Mississippi, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JVR CC PAF HT KS. Performed the experiments: JVR CC PAF HT KS. Analyzed the data: JVR CC PAF HT KS. Contributed reagents/materials/analysis tools: JVR CC PAF HT KS. Wrote the paper: JVR CC PAF HT KS.