About the Authors

Chen-Ching Lin

Affiliation Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Ramkrishna Mitra

Affiliation Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Zhongming Zhao

zhongming.zhao@vanderbilt.edu

Affiliations Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Psychiatry, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Center for Quantitative Sciences, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Competing Interests

The corresponding author, Dr. Zhongming Zhao, is an editor of PLoS ONE. According to the PLOS ONE Editorial policies and criteria, the authors claim “this does not alter the authors’ adherence to PLOS ONE Editorial policies and criteria”.

Author Contributions

Conceived and designed the experiments: ZZ CCL. Performed the experiments: CCL RM. Analyzed the data: CCL RM. Contributed reagents/materials/analysis tools: CCL RM. Wrote the paper: ZZ CCL RM.