About the Authors

Laura M. Dixon

laura.dixon@sruc.ac.uk

Affiliation Animal and Veterinary Science, SRUC, Edinburgh, United Kingdom

Sarah Brocklehurst

Affiliation Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Vicky Sandilands

Affiliation Animal and Veterinary Science, SRUC, Edinburgh, United Kingdom

Melissa Bateson

Affiliation Center for Behaviour and Evolution, Institute of Neuroscience, Newcastle University, United Kingdom

Bert J. Tolkamp

Affiliation Animal and Veterinary Science, SRUC, Edinburgh, United Kingdom

Rick B. D'Eath

Affiliation Animal and Veterinary Science, SRUC, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LMD SB VS MB BJT RBD. Performed the experiments: LMD. Analyzed the data: SB. Wrote the paper: LMD SB RBD.