About the Authors

Taku Kadoya

kadoya@nies.go.jp

Affiliation Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, Onogawa, Tsukuba, Ibaraki, Japan

Akio Takenaka

Affiliation Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, Onogawa, Tsukuba, Ibaraki, Japan

Fumiko Ishihama

Affiliation Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, Onogawa, Tsukuba, Ibaraki, Japan

Taku Fujita

Affiliation Nature Conservation Society of Japan, Shinkawa, Chuo-ku, Tokyo, Japan

Makoto Ogawa

Affiliation Tokushima Prefectural Museum, Bunka-no-mori Park, Hachiman-cho, Tokushima-shi, Tokushima, Japan

Teruo Katsuyama

Affiliation Kanagawa Prefectural Museum of Natural History, Odawara, Kanagawa, Japan

Yasuro Kadono

Affiliation Faculty of Science, Kobe University, Kobe, Japan

Nobumitsu Kawakubo

Affiliation Faculty of Applied Biological Sciences, Gifu University, Yanagido, Gifu, Japan

Shunsuke Serizawa

Affiliation Department of Biology, Aichi Kyoiku University, Igaya-cho, Kariya, Aichi, Japan

Hideki Takahashi

Affiliation Hokkaido University Museum, Sapporo, Hokkaido, Japan

Masayuki Takamiya

Affiliation Department of Environmental Science, Kumamoto University, Kumamoto, Japan

Shinji Fujii

Affiliation Division of Human Environment, University of Human Environments, Okazaki, Aichi, Japan

Hiroyuki Matsuda

Affiliation Faculty of Natural Environment and Information Sciences, Yokohama National University, Tokiwadai, Hodogaya-ku, Yokohama city, Kanagawa, Japan

Kazuo Muneda

Affiliation Faculty of Natural Environment and Information Sciences, Yokohama National University, Tokiwadai, Hodogaya-ku, Yokohama city, Kanagawa, Japan

Masatsugu Yokota

Affiliation Faculty of Science, University of the Ryukyus, Senbaru, Nishihara, Nakagami, Okinawa, Japan

Koji Yonekura

Affiliation Center for Academic Resources and Archives, Botanical Gardens, Tohoku University, Aoba, Aramaki, Aoba-ku, Sendai, Miyagi, Japan

Tetsukazu Yahara

Affiliation Department of Biology, Kyushu University, Higashi-ku, Fukuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK TY. Analyzed the data: TK FI AT. Contributed reagents/materials/analysis tools: TK FI AT TF MO TK YK NK SS HT MT SF HM KM MY KY TY. Wrote the paper: TK FI AT TY.