About the Authors

Alistair C. Darby

Contributed equally to this work with: Alistair C. Darby, Colin J. McInnes

Affiliation University of Liverpool, Liverpool, United Kingdom

Colin J. McInnes

Contributed equally to this work with: Alistair C. Darby, Colin J. McInnes

Affiliation Moredun Research Institute, Penicuik, United Kingdom

Karina Hansen Kjær

Affiliation University of Aarhus, Aarhus, Denmark

Ann R. Wood

Affiliation Moredun Research Institute, Penicuik, United Kingdom

Margaret Hughes

Affiliation University of Liverpool, Liverpool, United Kingdom

Pia Møller Martensen

Affiliation University of Aarhus, Aarhus, Denmark

Alan D. Radford

Affiliation University of Liverpool, Liverpool, United Kingdom

Neil Hall

Neil.Hall@liv.ac.uk

Affiliations University of Liverpool, Liverpool, United Kingdom, King Abdulaziz University, Jeddah, Saudi Arabia

Julian Chantrey

Affiliation University of Liverpool, Liverpool, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ACD CJM ADR JC NH PMM. Performed the experiments: MH ARW KHK. Analyzed the data: NH ACD ADR CJM KHK. Contributed reagents/materials/analysis tools: CJM. Wrote the paper: CJM ACD AR NH JC.