About the Authors

Skye A. Geherin

Contributed equally to this work with: Skye A. Geherin, R. Paul Wilson, Silke Jennrich

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

R. Paul Wilson

Contributed equally to this work with: Skye A. Geherin, R. Paul Wilson, Silke Jennrich

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Silke Jennrich

Contributed equally to this work with: Skye A. Geherin, R. Paul Wilson, Silke Jennrich

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Gudrun F. Debes

gdebes@vet.upenn.edu

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GFD SAG SJ RPW. Performed the experiments: GFD SAG SJ RPW. Analyzed the data: GFD SAG SJ RPW. Wrote the paper: GFD SAG SJ RPW.