About the Authors

John P. Clarkson

john.clarkson@warwick.ac.uk

Affiliation Warwick Crop Centre, School of Life Sciences, University of Warwick, Wellesbourne, Warwick, United Kingdom

Laura Fawcett

Affiliation ADAS UK Ltd, Pendeford House, Pendeford, Wolverhampton, United Kingdom

Steven G. Anthony

Affiliation ADAS UK Ltd, Pendeford House, Pendeford, Wolverhampton, United Kingdom

Caroline Young

Affiliation ADAS UK Ltd, ADAS Drayton, Defra Drayton, Stratford upon Avon, United Kingdom

Competing Interests

LF, SGA and CY are employees of ADAS UK Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JPC LF SA CY. Performed the experiments: JPC. Analyzed the data: JPC LF SGA. Contributed reagents/materials/analysis tools: JPC LF SGA. Wrote the paper: JC LF CY.