About the Authors

Liyan He

Affiliations Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, University of Chinese Academy of Sciences, Beijing, China

Xiaotian Han

Affiliation Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Zhiming Yu

zyu@qdio.ac.cn

Affiliation Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LH XH ZY. Performed the experiments: LH. Analyzed the data: LH XH ZY. Contributed reagents/materials/analysis tools: XH ZY. Wrote the paper: LH.