About the Authors

Paige A. Winkler

Affiliations Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America, Genetics Program, Michigan State University, East Lansing, Michigan, United States of America

Kara R. Gornik

Affiliation Cummings School of Veterinary Medicine, Tufts University, North Grafton, Massachusetts, United States of America

David T. Ramsey

Affiliation The Animal Ophthalmology Center, Williamston, Michigan, United States of America

Richard R. Dubielzig

Affiliation Comparative Ocular Pathology Laboratory of Wisconsin, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Patrick J. Venta

Affiliations Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America, Genetics Program, Michigan State University, East Lansing, Michigan, United States of America

Simon M. Petersen-Jones

Affiliations Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America, Genetics Program, Michigan State University, East Lansing, Michigan, United States of America

Joshua T. Bartoe

bartoejo@cvm.msu.edu

Affiliation Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PAW DTR PJV SMP-J JTB. Performed the experiments: PAW KRG DTR RRD JTB. Analyzed the data: PAW RRD PJV SMP-J JTB. Wrote the paper: PAW PJV JTB.