About the Authors

Elizabeth M. Samuelson

Contributed equally to this work with: Elizabeth M. Samuelson, Renee M. Laird

Current address: Benaroya Research Institute, Seattle, Washington, United States of America

Affiliation Department of Microbiology and Immunology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Renee M. Laird

Contributed equally to this work with: Elizabeth M. Samuelson, Renee M. Laird

Current address: Naval Medical Research Center, Silver Spring, Maryland, United States of America

Affiliation Department of Microbiology and Immunology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Amber M. Papillion

Affiliation Department of Microbiology and Immunology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Arthur H. Tatum

Affiliation Department of Pathology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Michael F. Princiotta

Affiliation Department of Microbiology and Immunology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Sandra M. Hayes

HayesSa@upstate.edu

Affiliation Department of Microbiology and Immunology, State University of New York Upstate Medical University, Syracuse, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMH EMS RML. Performed the experiments: EMS RML SMH AMP MFP. Analyzed the data: EMS RML SMH AMP MFP AHT. Contributed reagents/materials/analysis tools: MFP. Wrote the paper: SMH.