About the Authors

Katherine E. Davis

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Sarah Fox

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

John Gigg

j.gigg@manchester.ac.uk

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Competing Interests

The authors have the following interests. JG is an in vivo electrophysiology consultant for GlaxoSmithKline Neurodegeneration, China. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KED JG. Performed the experiments: KED. Analyzed the data: KED SF JG. Contributed reagents/materials/analysis tools: SF. Wrote the paper: KED SF JG.