About the Authors

Sandra McCutcheon

sandra.mccutcheon@roslin.ed.ac.uk

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Jan P. M. Langeveld

Affiliation Central Veterinary Institute of Wageningen UR, Lelystad, The Netherlands

Boon Chin Tan

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Andrew C. Gill

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Christopher de Wolf

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Stuart Martin

Affiliation Animal Health and Veterinary Laboratories Agency, Lasswade Laboratory, Edinburgh, Scotland, United Kingdom

Lorenzo Gonzalez

Affiliation Animal Health and Veterinary Laboratories Agency, Lasswade Laboratory, Edinburgh, Scotland, United Kingdom

James Alibhai

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

A. Richard Alejo Blanco

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Lauren Campbell

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Nora Hunter

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

E. Fiona Houston

Affiliation Neurobiology Division, The Roslin Institute and Royal (Dick) School of Veterinary Sciences, Easter Bush, Edinburgh, Scotland, United Kingdom

Competing Interests

Dr. A. C. Gill is also an academic editor for PLOS ONE. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: S. McCutcheon JPML ACG LG JA LC NH EFH. Performed the experiments: S. McCutcheon JPML BCT ACG CdW S. Martin JA ARAB LC. Analyzed the data: S. McCutcheon JPML ACG S. Martin LG JA ARAB LC EFH. Contributed reagents/materials/analysis tools: S. McCutcheon JPML ACG S. Martin LG JA NH EFH. Wrote the paper: S. McCutcheon JPML ACG LG EFH.