About the Authors

Adam L. Bailey

Contributed equally to this work with: Adam L. Bailey, Michael Lauck

Affiliations Department of Pathology and Laboratory Medicine, University of Wisconsin–Madison, Madison, Wisconsin, United States of America, Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America

Michael Lauck

Contributed equally to this work with: Adam L. Bailey, Michael Lauck

Affiliations Department of Pathology and Laboratory Medicine, University of Wisconsin–Madison, Madison, Wisconsin, United States of America, Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America

Andrea Weiler

Affiliations Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Samuel D. Sibley

Affiliations Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Jorge M. Dinis

Affiliation Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Zachary Bergman

Affiliations Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Chase W. Nelson

Affiliation Department of Biological Sciences, University of South Carolina, Columbia, South Carolina, United States of America

Michael Correll

Affiliation Department of Computer Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Michael Gleicher

Affiliation Department of Computer Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

David Hyeroba

Affiliation Makerere University, Kampala, Uganda

Alex Tumukunde

Affiliation Makerere University, Kampala, Uganda

Geoffrey Weny

Affiliation Makerere University, Kampala, Uganda

Colin Chapman

Affiliations Makerere University, Kampala, Uganda, Department of Anthropology and School of Environment, McGill University, Montreal, Quebec, Canada

Jens H. Kuhn

Affiliation Integrated Research Facility at Fort Detrick, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Fort Detrick, Frederick, Maryland, United States of America

Austin L. Hughes

Affiliation Department of Biological Sciences, University of South Carolina, Columbia, South Carolina, United States of America

Thomas C. Friedrich

Affiliations Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Tony L. Goldberg

Affiliations Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America, Department of Pathobiological Sciences, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

David H. O'Connor

dhoconno@wisc.edu

Affiliations Department of Pathology and Laboratory Medicine, University of Wisconsin–Madison, Madison, Wisconsin, United States of America, Wisconsin National Primate Research Center, Madison, Wisconsin, United States of America

Competing Interests

Dr. Jens H. Kuhn is employed by a commercial company, Tunnell Government Services, Inc. Tunnell is a subcontractor to Battelle Memorial Institute, which is the primary contractor to NIH to run the NIH facility “Integrated Research Facility at Fort Detrick”. As the authors pointed out in the disclaimer, Tunnell's involvement with Battelle, and Battelle's involvement with NIH is stipulated in US Department of Health and Human Services contract HHSN272200700016I. All work performed by Jens H. Kuhn (and all other Tunnell IRF employees) is property of the US government under the contract. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ALB ML MG CC ALH TCF TLG DHO. Performed the experiments: ALB ML AW ZB CWN MC DH AT GW ALH. Analyzed the data: ALB ML SS JD ALH TCF TLG DHO. Contributed reagents/materials/analysis tools: ALB ML CWN MC MG DH AT GW ALH TCF TLG DHO. Wrote the paper: ALB JHK ALH TCF TLG DHO.