About the Authors

He Meng

Contributed equally to this work with: He Meng, Yan Zhang

menghe@sjtu.edu.cn (HM); yzhang@vbi.vt.edu (YZ); pbsiegel@vt.edu (PBS)

Affiliation Shanghai Key Laboratory of Veterinary Biotechnology, School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, P. R. China

Yan Zhang

Contributed equally to this work with: He Meng, Yan Zhang

menghe@sjtu.edu.cn (HM); yzhang@vbi.vt.edu (YZ); pbsiegel@vt.edu (PBS)

Affiliation Virginia Bioinformatics Institute, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Lele Zhao

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, P. R. China

Wenjing Zhao

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, P. R. China

Chuan He

Affiliation Shanghai Personal Biotechnology Limited Company, Shanghai, P. R. China

Christa F. Honaker

Affiliation Department of Animal and Poultry Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Zhengxiao Zhai

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, P. R. China

Zikui Sun

Affiliation Shanghai Personal Biotechnology Limited Company, Shanghai, P. R. China

Paul B. Siegel

menghe@sjtu.edu.cn (HM); yzhang@vbi.vt.edu (YZ); pbsiegel@vt.edu (PBS)

Affiliation Department of Animal and Poultry Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Competing Interests

Chuan He and Zikui Sun are employed by Shanghai Personal Biotechnology Limited Company. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HM PBS YZ. Performed the experiments: LZ WZ YZ ZZ ZS. Analyzed the data: CH LZ WZ YZ. Contributed reagents/materials/analysis tools: CFH PBS YZ. Wrote the manuscript: CFH HM PBS YZ.