About the Authors

Tarja Joensuu

Contributed equally to this work with: Tarja Joensuu, Saara Tegelberg

tarja.joensuu@helsinki.fi

Affiliations Folkhälsan Institute of Genetics, Helsinki, Finland, Department of Medical Genetics, Haartman Institute and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, Neuroscience Center, University of Helsinki, Helsinki, Finland

Saara Tegelberg

Contributed equally to this work with: Tarja Joensuu, Saara Tegelberg

Affiliations Folkhälsan Institute of Genetics, Helsinki, Finland, Department of Medical Genetics, Haartman Institute and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, Neuroscience Center, University of Helsinki, Helsinki, Finland

Eva Reinmaa

Current address: Department of Biotechnology, Institute of Molecular and Cell Biology and Estonian Genome Center, University of Tartu, Tartu, Estonia

Affiliation Folkhälsan Institute of Genetics, Helsinki, Finland

Mikael Segerstråle

Affiliations Neuroscience Center, University of Helsinki, Helsinki, Finland, Department of Biosciences, Faculty of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, Faculty of Veterinary Medicine, University of Helsinki, Helsinki, Finland

Paula Hakala

Affiliations Folkhälsan Institute of Genetics, Helsinki, Finland, Department of Medical Genetics, Haartman Institute and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, Neuroscience Center, University of Helsinki, Helsinki, Finland

Heidi Pehkonen

Affiliation Institute of Biomedicine, Pharmacology, University of Helsinki, Helsinki, Finland

Esa R. Korpi

Affiliation Institute of Biomedicine, Pharmacology, University of Helsinki, Helsinki, Finland

Jaana Tyynelä

Affiliation Institute of Biomedicine, Biochemistry and Developmental Biology, University of Helsinki, Helsinki, Finland

Tomi Taira

Affiliations Neuroscience Center, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, Faculty of Veterinary Medicine, University of Helsinki, Helsinki, Finland

Iiris Hovatta

Affiliations Department of Biosciences, Faculty of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland, Mental Health and Substance Abuse Services, National Institute for Health and Welfare, Helsinki, Finland

Outi Kopra

Affiliations Folkhälsan Institute of Genetics, Helsinki, Finland, Department of Medical Genetics, Haartman Institute and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, Neuroscience Center, University of Helsinki, Helsinki, Finland

Anna-Elina Lehesjoki

Affiliations Folkhälsan Institute of Genetics, Helsinki, Finland, Department of Medical Genetics, Haartman Institute and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, Neuroscience Center, University of Helsinki, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TJ ST ERK TT OK A-EL. Performed the experiments: TJ ST ER MS PH HP OK. Analyzed the data: TJ ST ER MS ERK JT TT IH OK. Contributed reagents/materials/analysis tools: ERK JT TT A-EL. Wrote the paper: TJ ST MS ERK JT TT IH OK A-EL.