About the Authors

Clio Der Sarkissian

clio.dersarkissian@gmail.com (CDS); wolfgang.haak@adelaide.edu.au (WH)

Current address: Centre for Geogenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Paul Brotherton

Current address: Archaeogenetics Research Group, School of Applied Sciences, University of Huddersfield, Huddersfield, United Kingdom

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Oleg Balanovsky

Affiliations Vavilov Institute for General Genetics, Russian Academy of Sciences, Moscow, Russia, Research Centre for Medical Genetics, Russian Academy of Medical Sciences, Moscow, Russia

Jennifer E. L. Templeton

Current address: School of Biological Sciences, Flinders University, Adelaide, South Australia, Australia

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Bastien Llamas

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Julien Soubrier

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Vyacheslav Moiseyev

Affiliation Peter the Great Museum of Anthropology and Ethnography (Kunstkamera) RAS, St Petersburg, Russia

Valery Khartanovich

Affiliation Peter the Great Museum of Anthropology and Ethnography (Kunstkamera) RAS, St Petersburg, Russia

Alan Cooper

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

Wolfgang Haak

clio.dersarkissian@gmail.com (CDS); wolfgang.haak@adelaide.edu.au (WH)

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia, Australia

The Genographic Consortium

Membership of The Genographic Consortium is provided in the Acknowledgments.

Competing Interests

The authors declare having received funding from a commercial partner (IBM). This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CDS WH PB BL AC. Performed the experiments: CDS WH PB BL JT. Analyzed the data: CDS WH PB BL JS. Contributed reagents/materials/analysis tools: CDS WH PB BL JS JT OB VM VK AC TGC. Wrote the paper: CDS WH AC.