About the Authors

Hirokazu Toju

toju.hirokazu.4c@kyoto-u.ac.jp

Affiliations Graduate School of Global Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan, Graduate School of Human and Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan

Hirotoshi Sato

Affiliations Graduate School of Global Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan, Graduate School of Human and Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan

Akifumi S. Tanabe

Affiliations Graduate School of Global Environmental Studies, Kyoto University, Sakyo, Kyoto, Japan, National Research Institute of Fisheries Science, Fisheries Research Agency, Fukuura, Kanazawa, Yokohama, Kanagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HT HS. Performed the experiments: HT HS. Analyzed the data: HT AST. Contributed reagents/materials/analysis tools: HT HS. Wrote the paper: HT HS AST.