About the Authors

Philip L. Reno

Contributed equally to this work with: Philip L. Reno, Cory Y. McLean

philreno@psu.edu (PLR); cmclean@stanfordalumni.org (CYM)

Affiliation Department of Anthropology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Cory Y. McLean

Contributed equally to this work with: Philip L. Reno, Cory Y. McLean

philreno@psu.edu (PLR); cmclean@stanfordalumni.org (CYM)

Current address: 23 and Me, Inc., Mountain View, California, United States of America

Affiliation Department of Computer Science, Stanford University, Stanford, California, United States of America

Jasmine E. Hines

Affiliation Department of Anthropology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Terence D. Capellini

Current address: Department of Human Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Gill Bejerano

Affiliations Department of Computer Science, Stanford University, Stanford, California, United States of America, Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

David M. Kingsley

Affiliations Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America, Howard Hughes Medical Institute, Stanford, California, United States of America

Competing Interests

The authors have the following interests. G.B. is a Microsoft Research Fellow. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: PLR CYM GB DMK. Performed the experiments: PLR CYM JEH TDC. Analyzed the data: PLR CYM JEH TDC GB DMK. Wrote the manuscript: PLR CYM DMK GB JEH TDC.