About the Authors

Yong Hun Jo

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Bharat Bhusan Patnaik

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Se Won Kang

Affiliation Department of Life Science and Biotechnology, College of Natural Sciences, Soonchunhyang University, Asan, South Korea

Sung-Hwa Chae

Affiliation GnC Bio Company Limited, Daejeon, South Korea

Seunghan Oh

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Dong Hyun Kim

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Mi Young Noh

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Gi Won Seo

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Heon Cheon Jeong

Affiliation Hampyeong County Insect Institute, Hampyeong County Agricultural Technology Center, Hampyeong, South Korea

Ju Young Noh

Affiliation Hampyeong County Insect Institute, Hampyeong County Agricultural Technology Center, Hampyeong, South Korea

Ji Eun Jeong

Affiliation Department of Life Science and Biotechnology, College of Natural Sciences, Soonchunhyang University, Asan, South Korea

Hee Ju Hwang

Affiliation Department of Life Science and Biotechnology, College of Natural Sciences, Soonchunhyang University, Asan, South Korea

Kisung Ko

Affiliation Department of Medicine, Medical Research Institute, College of Medicine, Chung-Ang University, Seoul, South Korea

Yeon Soo Han

Affiliation Division of Plant Biotechnology, College of Agriculture and Life Sciences, Chonnam National University, Gwangju, South Korea

Yong Seok Lee

yslee@sch.ac.kr

Affiliation Department of Life Science and Biotechnology, College of Natural Sciences, Soonchunhyang University, Asan, South Korea

Competing Interests

Dr. Sung-Hwa Chae, who is currently working at GnC Bio Co., Ltd., 4F, Yekun-plaza, 621-6, Banseok-dong, Yuseong-gu, Daejeon 305–150, South Korea, is one of the co-authors for the manuscript. Dr. Sung-Hwa Chae does not hold any competing interests on the publication of the manuscript, consultancy, patents, products in development or marketed products. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YSL YSH BBP. Performed the experiments: YHJ SWK SHC MYN GWS HJH. Analyzed the data: BBP SO HCJ JYN JEJ KK. Contributed reagents/materials/analysis tools: YSL YSH BBP DHK. Wrote the paper: BBP YSL YSH KK.