About the Authors

Natalie Jiricny

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Søren Molin

Affiliation Department of Systems Biology, Technical University of Denmark, Lyngby, Denmark

Kevin Foster

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Stephen P. Diggle

Affiliation School of Life Sciences, Centre for Biomolecular Sciences, University Park, University of Nottingham, Nottingham, United Kingdom

Pauline D. Scanlan

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Melanie Ghoul

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Helle Krogh Johansen

Affiliation Institute for International Health, Immunology and Microbiology, University of Copenhagen, Copenhagen, Denmark

Lorenzo A. Santorelli

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Roman Popat

Affiliation Centre for Immunity, Infection and Evolution, University of Edinburgh Ashworth Labs, King's Buildings, West Mains Road, Edinburgh, United Kingdom

Stuart A. West

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Ashleigh S. Griffin

ashleigh.griffin@zoo.ox.ac.uk

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Competing Interests

The authors would like to declare here that this work was supported in part by a commercial funder - L'Oreal - through a Fellowship for Women in Science to Ashleigh Griffin in 2008. This does not alter the authors′ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NJ KF SPD SAW ASG. Performed the experiments: NJ PDS MG LAS RP. Analyzed the data: NJ MG. Contributed reagents/materials/analysis tools: SM KF SPD HKJ SAW ASG. Wrote the paper: NJ SM KF SPD PDS MG HKJ LAS RP SAW ASG.