About the Authors

Li Li

Contributed equally to this work with: Li Li, Delin Li

Affiliations College of Agronomy, Northwest Agriculture & Forestry University, Yangling, Shaanxi, China, Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Delin Li

Contributed equally to this work with: Li Li, Delin Li

Current address: DATA Biotechnology Beijing limited company, Beijing, Hebei, China

Affiliation Department of Plant Genetics & Breeding, China Agricultural University, Beijing, Hebei, China

Sanzhen Liu

Current address: Department of Plant Pathology, Kansas State University, Manhattan, Kansas, USA

Affiliation Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Xiaoli Ma

Affiliation Department of Plant Genetics & Breeding, China Agricultural University, Beijing, Hebei, China

Charles R. Dietrich

Current address: Monsanto, Chesterfield, Missouri, USA

Affiliation Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Heng-Cheng Hu

Affiliation Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Gaisheng Zhang

Affiliation College of Agronomy, Northwest Agriculture & Forestry University, Yangling, Shaanxi, China

Zhiyong Liu

Affiliation Department of Plant Genetics & Breeding, China Agricultural University, Beijing, Hebei, China

Jun Zheng

Affiliation Institute of Crop Sciences, Chinese Academy of Agricultural Sciences, Beijing, Hebei, China

Guoying Wang

Affiliation Institute of Crop Sciences, Chinese Academy of Agricultural Sciences, Beijing, Hebei, China

Patrick S. Schnable

Schnable@iastate.edu

Affiliations Department of Agronomy, Iowa State University, Ames, Iowa, United States of America, Department of Plant Genetics & Breeding, China Agricultural University, Beijing, Hebei, China, Center for Plant Genomics, Iowa State University, Ames, Iowa, United States of America

Competing Interests

Although co-author Delin Li is currently employed by DATA Biotechnology Beijing limited company all of his contributions to the current manuscript were completed while he was a graduate student at Chinese Agricultural University (CUA). A similar situation affects author Chuck Dietrich (i.e., although he is currently employed by Monsanto, the research reported in this manuscript was conducted while he was a student at Iowa State University). Thus, the authors do not have any conflicts of interest to report here.

Author Contributions

Conceived and designed the experiments: SL PS. Performed the experiments: LL DL SL CD XM. Analyzed the data: SL DL. Contributed reagents/materials/analysis tools: JZ GW. Wrote the manuscript: LL HH PS. Mentored graduate student authors: GZ ZL PS.