About the Authors

Suzanne J. Rowe

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Scotland, United Kingdom

Amy Rowlatt

Affiliation The Roslin Institute, The University of Edinburgh, Roslin, Scotland, United Kingdom

Gail Davies

Affiliation Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Sarah E. Harris

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom, Medical Genetics Section, The University of Edinburgh, Edinburgh, Scotland, United Kingdom

David J. Porteous

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom, Medical Genetics Section, The University of Edinburgh, Edinburgh, Scotland, United Kingdom

David C. Liewald

Affiliation Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Geraldine McNeill

Affiliation Institute of Applied Health Sciences, University of Aberdeen, Aberdeen, Scotland, United Kingdom

John M. Starr

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom, Alzheimer Scotland Dementia Research Centre, The University of Edinburgh, Edinburgh, Scotland, United Kingdom

Ian J. Deary

Contributed equally to this work with: Ian J. Deary, Albert Tenesa

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, Scotland, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Albert Tenesa

Contributed equally to this work with: Ian J. Deary, Albert Tenesa

Albert.Tenesa@ed.ac.uk

Affiliations The Roslin Institute, The University of Edinburgh, Roslin, Scotland, United Kingdom, Medical Research Council Human Genetics Unit at the Medical Research Council Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AT. Performed the experiments: AT ID SR JS. Analyzed the data: SR AT AR. Contributed reagents/materials/analysis tools: GD SH DP DL GM JS ID AT. Wrote the paper: SR AT. Critical read of manuscript: SR GD SH DP DL GM JS ID AT AR.