About the Authors

Yohei Nakamura

ynakamura@kochi-u.ac.jp

Affiliation Graduate School of Kuroshio Science, Kochi University, Nankoku, Kochi, Japan

David A. Feary

Affiliation School of the Environment, University of Technology, Sydney, New South Wales, Australia

Masaru Kanda

Affiliations Graduate School of Kuroshio Science, Kochi University, Nankoku, Kochi, Japan, Kuroshio Zikkan Center, Otsuki, Kochi, Japan

Kosaku Yamaoka

Affiliation Graduate School of Kuroshio Science, Kochi University, Nankoku, Kochi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN. Performed the experiments: YN MK KY. Analyzed the data: YN DAF. Wrote the paper: YN DAF.