About the Authors

James J. Burston

Contributed equally to this work with: James J. Burston, Devi Rani Sagar

Affiliations Arthritis Research UK Pain Centre, University of Nottingham, Nottingham, United Kingdom, School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Devi Rani Sagar

Contributed equally to this work with: James J. Burston, Devi Rani Sagar

Affiliations Arthritis Research UK Pain Centre, University of Nottingham, Nottingham, United Kingdom, School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Pin Shao

Affiliation Department of Radiology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Mingfeng Bai

Affiliations Department of Radiology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, University of Pittsburgh Cancer Institute, Pittsburgh, Pennsylvania, United States of America

Emma King

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Louis Brailsford

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Jenna M. Turner

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Gareth J. Hathway

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Andrew J. Bennett

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

David A. Walsh

Affiliation Arthritis Research UK Pain Centre, University of Nottingham, Nottingham, United Kingdom

David A. Kendall

Affiliation School of Life Sciences, University of Nottingham, Nottingham, United Kingdom

Aron Lichtman

Affiliation Department of Pharmacology and Toxicology, Virginia Commonwealth University, Richmond, Virginia, United States of America

Victoria Chapman

victoria.chapman@nottingham.ac.uk

Affiliation Arthritis Research UK Pain Centre, University of Nottingham, Nottingham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DRS JJB AJB DAK GJH DAW VC. Performed the experiments: DRS JJB LB JMT EK PS MB. Analyzed the data: DRS JJB LB JMT AB GJH VC. Contributed reagents/materials/analysis tools: PS MB AL. Wrote the paper: DRS JJB PS MB EK GJH AJB DAW DAK AL VC.