About the Authors

Kunitake Higo

Affiliation Research Institute for Biomedical Sciences, Tokyo University of Science, Noda-shi, Chiba, Japan

Teikichi Ikura

Affiliation Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Masayuki Oda

Affiliation Graduate School of Life and Environmental Sciences, Kyoto Prefectural University, Sakyo-ku, Kyoto-shi, Kyoto, Japan

Hisayuki Morii

Affiliation Biomedical Research Institute, National Institute of Advanced Industrial Science and Technology, Tsukuba-shi, Ibaraki, Japan

Jun Takahashi

Affiliation Research Institute for Biomedical Sciences, Tokyo University of Science, Noda-shi, Chiba, Japan

Ryo Abe

Affiliation Research Institute for Biomedical Sciences, Tokyo University of Science, Noda-shi, Chiba, Japan

Nobutoshi Ito

ito.str@tmd.ac.jp

Affiliation Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NI RA MO. Performed the experiments: KH TI JT NI. Analyzed the data: NI HM. Contributed reagents/materials/analysis tools: HM. Wrote the paper: NI KH MO.