About the Authors

Zeng Wang

Affiliation Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, School of Life Sciences, Xiamen University, Xiamen, Fujian, People’s Republic of China

Xiaoping Zhou

Affiliations Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, School of Life Sciences, Xiamen University, Xiamen, Fujian, People’s Republic of China, Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, College of the Environment and Ecology, Xiamen University, Xiamen, Fujian, People’s Republic of China

Qingxian Lin

Affiliations Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, School of Life Sciences, Xiamen University, Xiamen, Fujian, People’s Republic of China, Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, College of the Environment and Ecology, Xiamen University, Xiamen, Fujian, People’s Republic of China

Wenzhen Fang

Contributed equally to this work with: Wenzhen Fang, Xiaolin Chen

wzfang@xmu.edu.cn (WF); xlchen@xmu.edu.cn (XC)

Affiliations Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, School of Life Sciences, Xiamen University, Xiamen, Fujian, People’s Republic of China, Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, College of the Environment and Ecology, Xiamen University, Xiamen, Fujian, People’s Republic of China

Xiaolin Chen

Contributed equally to this work with: Wenzhen Fang, Xiaolin Chen

wzfang@xmu.edu.cn (WF); xlchen@xmu.edu.cn (XC)

Affiliations Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, School of Life Sciences, Xiamen University, Xiamen, Fujian, People’s Republic of China, Key Laboratory of Ministry of Education for Coast and Wetland Ecosystems, College of the Environment and Ecology, Xiamen University, Xiamen, Fujian, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XC ZW WF. Performed the experiments: ZW QL. Analyzed the data: ZW XZ XC WF. Contributed reagents/materials/analysis tools: XC WF XZ. Wrote the manuscript: ZW XZ XC WF. Edited manuscript: XC WF. Graduate Supervisor: XC WF. Funding Source: XC WF XZ.