About the Authors

Cui Guan

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, P.R. of China

Andrew B. Barron

Affiliation Department of Biological Sciences, Macquarie University, North Ryde, New South Wales, Australia

Xu Jiang He

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, P.R. of China

Zi Long Wang

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, P.R. of China

Wei Yu Yan

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, P.R. of China

Zhi Jiang Zeng

bees1965@sina.com

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, P.R. of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZJZ CG. Performed the experiments: CG. Analyzed the data: CG ABB ZLW. Contributed reagents/materials/analysis tools: XJH WYY. Wrote the paper: CG ABB ZJZ. Approved the manuscript: CG ABB XJH ZLW WYY ZJZ.