About the Authors

Kirsty Kemmett

K.Kemmett@liv.ac.uk

Affiliation Institute of Infection and Global Health, Department of Infection Biology, School of Veterinary Science, The University of Liverpool, Neston, Cheshire, United Kingdom

Tom Humphrey

Affiliation Institute of Infection and Global Health, Department of Infection Biology, School of Veterinary Science, The University of Liverpool, Neston, Cheshire, United Kingdom

Steven Rushton

Affiliation School of Biology, Newcastle University, Newcastle Upon Tyne, United Kingdom

Andrew Close

Affiliation School of Biology, Newcastle University, Newcastle Upon Tyne, United Kingdom

Paul Wigley

Affiliation Institute of Infection and Global Health, Department of Infection Biology, School of Veterinary Science, The University of Liverpool, Neston, Cheshire, United Kingdom

Nicola J. Williams

Affiliation Institute of Infection and Global Health, Department of Epidemiology and Population Health, School of Veterinary Science, The University of Liverpool, Neston, Cheshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KK NJW TH PW. Performed the experiments: KK. Analyzed the data: KK SR AC NJW. Contributed reagents/materials/analysis tools: NJW TH AC SR. Wrote the paper: KK NJW SR PW TH.