About the Authors

Ryoji Shinya

shinya@caltech.edu

Affiliations Graduate School of Agriculture, Kyoto University, Kyoto, Japan, College of Bioscience and Biotechnology, Chubu University, Kasugai, Japan

Hironobu Morisaka

Contributed equally to this work with: Hironobu Morisaka, Taisei Kikuchi

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Taisei Kikuchi

Contributed equally to this work with: Hironobu Morisaka, Taisei Kikuchi

Affiliation Forestry and Forest Products Research Institute, Tsukuba, Japan

Yuko Takeuchi

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Mitsuyoshi Ueda

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Kazuyoshi Futai

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS HM MU KF. Performed the experiments: RS HM TK. Analyzed the data: RS HM TK. Contributed reagents/materials/analysis tools: RS HM TK MU. Wrote the paper: RS TK YT.