About the Authors

Mark Clendenning

Affiliation Cancer and Population Studies Group, Queensland Institute of Medical Research, Brisbane, Queensland, Australia

Joanne P. Young

Affiliation Cancer and Population Studies Group, Queensland Institute of Medical Research, Brisbane, Queensland, Australia

Michael D. Walsh

Affiliations Cancer and Population Studies Group, Queensland Institute of Medical Research, Brisbane, Queensland, Australia, Department of Histopathology, Sullivan Nicolaides Pathology, Brisbane, Queensland, Australia

Sonja Woodall

Affiliation New Zealand Familial Gastrointestinal Cancer Service, Auckland Hospital, Auckland, New Zealand

Julie Arnold

Affiliation New Zealand Familial Gastrointestinal Cancer Service, Auckland Hospital, Auckland, New Zealand

Mark Jenkins

Affiliation Centre for MEGA Epidemiology, University of Melbourne, Melbourne, Victoria, Australia

Aung Ko Win

Affiliation Centre for MEGA Epidemiology, University of Melbourne, Melbourne, Victoria, Australia

John L. Hopper

Affiliation Centre for MEGA Epidemiology, University of Melbourne, Melbourne, Victoria, Australia

Kevin Sweet

Affiliation Division of Human Genetics, The Ohio State University Medical Centre, Columbus, Ohio, United States of America

Steven Gallinger

Affiliations Cancer Care Ontario, Toronto, Ontario, Canada, Samuel Lunenfeld Research Institute, Mount Sinai Hospital, Toronto, Ontario, Canada, Zane Cohen Centre for Digestive Diseases, Mount Sinai Hospital, Toronto, Ontario, Canada

Christophe Rosty

Affiliations Cancer and Population Studies Group, Queensland Institute of Medical Research, Brisbane, Queensland, Australia, Department of Molecular and Cellular Pathology, University of Queensland, Brisbane, Queensland, Australia, Envoi Specialist Pathologists, Brisbane, Queensland, Australia

Susan Parry

Affiliations New Zealand Familial Gastrointestinal Cancer Service, Auckland Hospital, Auckland, New Zealand, Department of Gastroenterology and Hepatology, Middlemore Hospital, Auckland, New Zealand

Daniel D. Buchanan

Daniel.Buchanan@qimr.edu.au

Affiliation Cancer and Population Studies Group, Queensland Institute of Medical Research, Brisbane, Queensland, Australia

Competing Interests

Dr. Michael D. Walsh is employed by Sullivan Nicolaides Pathology and Dr. Christophe Rosty is employed by Envoi Specialist Pathologists, both are private diagnostic pathology services. The authors have no conflicts of interest to declare with regard to their employment and this does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DDB JPY MC SP CR MDW KS AKW MJ JLH SG. Performed the experiments: MC. Analyzed the data: DDB MC CR JPY MDW SP KS MJ AKW JLH SG. Contributed reagents/materials/analysis tools: SP JA SW SG. Wrote the paper: DDB JPY MC MDW CR SP KS AKW MJ JLH SG SW JA.