About the Authors

Ching-Yu Lin

Contributed equally to this work with: Ching-Yu Lin, Chieh Huo, Li-Kuo Kuo

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, National Institute of Cancer Research, National Health Research Institutes, Miaoli, Taiwan

Chieh Huo

Contributed equally to this work with: Ching-Yu Lin, Chieh Huo, Li-Kuo Kuo

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, Department of Life Sciences, National Central University, Taoyuan, Taiwan

Li-Kuo Kuo

Contributed equally to this work with: Ching-Yu Lin, Chieh Huo, Li-Kuo Kuo

Affiliation Division of Pulmonary and Critical Care Medicine, Department of Internal Medicine, Mackay Memorial Hospital, Taipei, Taiwan

Richard A. Hiipakka

Affiliation Ben May Department for Cancer Research, The University of Chicago, Illinois, United States of America

Richard Baker Jones

Affiliations Ben May Department for Cancer Research, The University of Chicago, Illinois, United States of America, Institute for Genomics and Systems Biology, The University of Chicago, Illinois, United States of America

Hui-Ping Lin

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan

Yuwen Hung

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, Taiwan

Liang-Cheng Su

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan

Jen-Chih Tseng

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, Taiwan

Ying-Yu Kuo

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan

Yu-Ling Wang

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Institute of Molecular Medicine, National Tsing Hua University, Hsinchu, Taiwan

Yasuhisa Fukui

Affiliation Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan

Yung-Hsi Kao

Affiliation Department of Life Sciences, National Central University, Taoyuan, Taiwan

John M. Kokontis

Affiliation Ben May Department for Cancer Research, The University of Chicago, Illinois, United States of America

Chien-Chih Yeh

Affiliations Division of Colon and Rectal Surgery, Taoyuan Armed Forces General Hospital, Taoyuan, Taiwan, Department of Surgery, Taoyuan Armed Forces General Hospital, Taoyuan, Taiwan

Linyi Chen

Affiliation Institute of Molecular Medicine, National Tsing Hua University, Hsinchu, Taiwan

Shiaw-Der Yang

Affiliation Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, Taiwan

Hsiao-Hui Fu

Affiliation Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu, Taiwan

Ya-Wen Chen

Affiliations Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, National Institute of Cancer Research, National Health Research Institutes, Miaoli, Taiwan

Kelvin K. C. Tsai

Affiliations Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, National Institute of Cancer Research, National Health Research Institutes, Miaoli, Taiwan

Jang-Yang Chang

Affiliations Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, National Institute of Cancer Research, National Health Research Institutes, Miaoli, Taiwan

Chih-Pin Chuu

cpchuu@nhri.org.tw

Affiliations Institute of Cellular and System Medicine, National Health Research Institutes, Miaoli, Taiwan, Translational Center for Glandular Malignancies, National Health Research Institutes, Miaoli, Taiwan, Graduate Program for Aging, China Medical University, Taichung, Taiwan, Ph.D. Program in Tissue Engineering and Regenerative Medicine, National Chung Hsing University, Taichung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CPC. Performed the experiments: CYL CH LKK HPL YWH YLW LCS YYK JCT RAH JMK CPC. Analyzed the data: CPC YF RAH JMK RBJ CCY HHF YHK LC SDY YWC KKT JYC. Contributed reagents/materials/analysis tools: YF RAH JMK CCY YHK HHF SDY YWC KKT JYC. Wrote the paper: CPC RAH JMK RBJ YWC LC.