About the Authors

Chizu Tanikawa

Affiliation Laboratory of Molecular Medicine, Human Genome Center, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Keitaro Matsuo

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Aichi, Japan

Michiaki Kubo

Affiliation Center for Genomic Medicine, The Institute of Physical and Chemical Research (RIKEN), Kanagawa, Japan

Atsushi Takahashi

Affiliation Center for Genomic Medicine, The Institute of Physical and Chemical Research (RIKEN), Kanagawa, Japan

Hidemi Ito

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Aichi, Japan

Hideo Tanaka

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Aichi, Japan

Yasushi Yatabe

Affiliation Department of Pathology and Molecular Diagnostics, Aichi Cancer Center Research Institute, Aichi, Japan

Kenji Yamao

Affiliation Department of Gastroenterology, Aichi Cancer Center Research Institute, Aichi, Japan

Naoyuki Kamatani

Affiliation Center for Genomic Medicine, The Institute of Physical and Chemical Research (RIKEN), Kanagawa, Japan

Kazuo Tajima

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Aichi, Japan

Yusuke Nakamura

Affiliations Laboratory of Molecular Medicine, Human Genome Center, Institute of Medical Science, The University of Tokyo, Tokyo, Japan, Departments of Medicine and Surgery, and Center for Personalized Therapeutics, The University of Chicago, Chicago, Illinois, United States of America

Koichi Matsuda

koichima@ims.u-tokyo.ac.jp

Affiliation Laboratory of Molecular Medicine, Human Genome Center, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CT K. Matsuda YN. Performed the experiments: CT K. Matsuo MK. Analyzed the data: CT AT NK HI. Contributed reagents/materials/analysis tools: HT YY KT KY. Wrote the paper: CT YN K. Matsuda.