About the Authors

Tshering Penjor

Affiliations Department of Applied Biological Sciences, Saga University, Honjo, Saga, Japan, Renewable Natural Resources Research Centre Wengkhar, Mongar, Bhutan

Masashi Yamamoto

Contributed equally to this work with: Masashi Yamamoto, Yukio Nagano

yamasa@agri.kagoshima-u.ac.jp (MY); nagano@cc.saga-u.ac.jp (YN)

Affiliation Faculty of Agriculture, Kagoshima University, Korimoto, Kagoshima, Japan

Miki Uehara

Affiliation Department of Applied Biological Sciences, Saga University, Honjo, Saga, Japan

Manami Ide

Affiliation Department of Applied Biological Sciences, Saga University, Honjo, Saga, Japan

Natsumi Matsumoto

Affiliation Department of Applied Biological Sciences, Saga University, Honjo, Saga, Japan

Ryoji Matsumoto

Affiliation Department of Applied Biological Sciences, Saga University, Honjo, Saga, Japan

Yukio Nagano

Contributed equally to this work with: Masashi Yamamoto, Yukio Nagano

yamasa@agri.kagoshima-u.ac.jp (MY); nagano@cc.saga-u.ac.jp (YN)

Affiliation Analytical Research Center for Experimental Sciences, Saga University, Honjo, Saga, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TP MY RM YN. Performed the experiments: TP MU MI NM YN. Analyzed the data: TP MY RM YN. Wrote the paper: TP MY YN.