About the Authors

Yan-Jie Li

Contributed equally to this work with: Yan-Jie Li, Yi Fang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Yi Fang

Contributed equally to this work with: Yan-Jie Li, Yi Fang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Ya-Ru Fu

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Jin-Guang Huang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Chang-Ai Wu

cczheng@sdau.edu.cn (CZ); cawu@sdau.edu.cn (CW)

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Cheng-Chao Zheng

cczheng@sdau.edu.cn (CZ); cawu@sdau.edu.cn (CW)

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YL Y. Fang Y. Fu JH CW CZ. Performed the experiments: YL Y. Fang Y. Fu. Analyzed the data: YL Y. Fang. Contributed reagents/materials/analysis tools: YL Y. Fang Y. Fu JH CW CZ. Wrote the paper: YL Y. Fang CZ.