About the Authors

Paul D. Jepson

paul.jepson@ioz.ac.uk

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Robert Deaville

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Karina Acevedo-Whitehouse

Affiliation Unit for Basic and Applied Microbiology, School of Natural Sciences, Autonomous University of Queretaro, Queretaro, Mexico

James Barnett

Affiliation Animal Health and Veterinary Laboratories Agency Truro, Polwhele, Truro, Cornwall, United Kingdom

Andrew Brownlow

Affiliation Scottish Agricultural College, Inverness, United Kingdom

Robert L. Brownell Jr.

Affiliation NOAA Fisheries, Southwest Fisheries Science Center, Pacific Grove, California, United States of America

Frances C. Clare

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Nick Davison

Affiliations Animal Health and Veterinary Laboratories Agency Truro, Polwhele, Truro, Cornwall, United Kingdom, Scottish Agricultural College, Inverness, United Kingdom

Robin J. Law

Affiliation Centre for Environment, Fisheries and Aquaculture Science, Lowestoft, United Kingdom

Jan Loveridge

Affiliation Cornwall Wildlife Trust Marine Strandings Network, Five Acres, Allet, Truro, United Kingdom

Shaheed K. Macgregor

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Steven Morris

Affiliation Centre for Environment, Fisheries and Aquaculture Science, Weymouth, Dorset, United Kingdom

Sinéad Murphy

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Rod Penrose

Affiliation Marine Environmental Monitoring, Penwalk, Llechryd, Cardigan, Ceredigion, United Kingdom

Matthew W. Perkins

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Eunice Pinn

Affiliation Joint Nature Conservation Committee, Aberdeen, United Kingdom

Henrike Seibel

Affiliation Institute of Terrestrial and Aquatic Wildlife Research, University of Veterinary Medicine Hannover, Büsum, Germany

Ursula Siebert

Affiliation Institute of Terrestrial and Aquatic Wildlife Research, University of Veterinary Medicine Hannover, Büsum, Germany

Eva Sierra

Affiliation Histology and Pathology Unit, Institute for Animal Health, Veterinary School Montana Cardones-Arucas, University of Las Palmas de Gran Canaria, Gran Canaria-Spain

Victor Simpson

Affiliation Wildlife Veterinary Investigation Centre, Chacewater, Truro, Cornwall, United Kingdom

Mark L. Tasker

Affiliation Joint Nature Conservation Committee, Aberdeen, United Kingdom

Nick Tregenza

Affiliation Cornwall Wildlife Trust Marine Strandings Network, Five Acres, Allet, Truro, United Kingdom

Andrew A. Cunningham

Affiliation Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Antonio Fernández

Affiliation Histology and Pathology Unit, Institute for Animal Health, Veterinary School Montana Cardones-Arucas, University of Las Palmas de Gran Canaria, Gran Canaria-Spain

Competing Interests

All acoustic and nonacoustic data relating to the naval exercises in this investigation was provided exclusively by the Royal Navy (UK Ministry of Defence). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: PDJ RD KAW ND RJL S. Morris S. Murphy HS US ES AAC AF. Performed the experiments: PDJ RD KAW JB FCC ND RJL SKM S. Morris S. Murphy RP MWP HS US ES VS AF. Analyzed the data: PDJ RD KAW JB AB RLB FCC ND RJL JL SKM S. Morris S. Murphy RP MWP EP HS US ES VS MLT NT AAC AF. Contributed reagents/materials/analysis tools: PDJ RD KAW JB ND RJL SKM S. Morris S. Murphy RP MWP HS US ES VS AF. Wrote the paper: PDJ RD KAW JB AB RLB FCC ND RJL JL S. Morris S. Murphy RP MWP EP HS US ES VS MLT NT AAC AF.