About the Authors

Fan Ding

Affiliation Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America

Jia Yao

Affiliation Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America

Liqin Zhao

Affiliation Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America

Zisu Mao

Affiliation Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America

Shuhua Chen

Affiliation Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America

Roberta Diaz Brinton

rbrinton@usc.edu

Affiliations Department of Pharmacology and Pharmaceutical Sciences, School of Pharmacy, University of Southern California, Los Angeles, California, United States of America, Department of Neurology, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FD JY RDB. Performed the experiments: FD JY ZM SC RDB. Analyzed the data: FD JY LZ RDB. Contributed reagents/materials/analysis tools: RDB. Wrote the paper: FD JY LZ RDB.