About the Authors

Ismail Mahat Bashir

Affiliations Walter Reed Project, Kenya Medical Research Institute, Kisumu, Kenya, Division of Malaria Vaccine Development, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Nekoye Otsyula

Affiliation Walter Reed Project, Kenya Medical Research Institute, Kisumu, Kenya

George Awinda

Affiliation Walter Reed Project, Kenya Medical Research Institute, Kisumu, Kenya

Michele Spring

Affiliation Division of Malaria Vaccine Development, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Petra Schneider

Joint senior author.

Affiliation Institutes of Evolution, Immunology and Infection Research, University of Edinburgh, Edinburgh, United Kingdom

John Njenga Waitumbi

john.waitumbi@usamru-k.org

Joint senior author.

Affiliation Walter Reed Project, Kenya Medical Research Institute, Kisumu, Kenya

Competing Interests

The authors wish to declare that GlaxoSmithKline Biologicals (GSK) provided their proprietary adjuvant “AS01B”. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Critical review and edits to the manuscript: PS. Conceived and designed the experiments: JNW NO MS. Performed the experiments: IMB GA. Analyzed the data: JNW IMB PS. Wrote the paper: JNW IMB PS NO MS.