About the Authors

Alexandra Mavroeidi

Affiliation Musculoskeletal Research Programme, University of Aberdeen, Aberdeen, United Kingdom

Lorna Aucott

Affiliation Musculoskeletal Research Programme, University of Aberdeen, Aberdeen, United Kingdom

Alison J. Black

Affiliation Musculoskeletal Research Programme, University of Aberdeen, Aberdeen, United Kingdom

William D. Fraser

Affiliation Norwich Medical School, University of East Anglia, Norwich, United Kingdom

David M. Reid

Affiliation Musculoskeletal Research Programme, University of Aberdeen, Aberdeen, United Kingdom

Helen M. Macdonald

h.macdonald@abdn.ac.uk

Affiliation Musculoskeletal Research Programme, University of Aberdeen, Aberdeen, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Instrumental in setting up the Aberdeen Prospective Osteoporosis Screening Study from where the volunteers were recruited: DMR. Conceived and designed the experiments: HMM DMR. Performed the experiments: AM. Analyzed the data: AM LA HMM. Contributed reagents/materials/analysis tools: WDF LA AJB. Wrote the paper: AM HMM WDF LA AJB DMR.